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☐			Preliminary Proxy Statement

☐			Confidential, for Use of the Commission Only (as permitted by Rule 14a-6(e)(2))

x			Definitive Proxy Statement

☐			Definitive Additional Materials

☐			Soliciting Material under §240.14a-12


Evelo Biosciences, Inc.
(Name of Registrant as Specified In Its Charter)

(Name of Person(s) Filing Proxy Statement, if Other Than The Registrant)


x			No fee required.
☐			Fee paid previously with preliminary materials
☐			Fee computed on table in exhibit required by Item 25(b) per Exchange Act Rules 14a-6(i)(1) and 0-11


			Page
GENERAL INFORMATIO N ABOUT THE ANNUAL MEETING AND VOTI NG			1
PROPOSAL 1 — ELECTION OF DIRECTORS			7
Board Size and Structure			7
Current Directors and Terms			7
Nominees for Director			7
Information About Board Nominees and Current and Continuing Directors			7
Nominees for Election to Three-Year Terms Expiring at the 202 5 Annual Meeting			8
Class I Directors Whose Terms Expire at the 202 2 Annual Meetin g Not Standing for Re-election			9
Class I I Directors Whose Terms Expire at the 202 3 Annual Meeting			9
Class I I I Directors Whose Terms Expire at the 202 4 Annual Meeting			10
Board Diversity			11
Board Recommendation			11
PROPOSAL 2 - RATIFICATION OF APPOINTMENT OF INDEPENDENT REGISTERED PUBLIC ACCOUNTING FIRM			12
Appointment of Independent Registered Public Accounting Firm			12
Independent Registered Public Accounting Firm Fees			12
Pre-Approval Policies and Procedures			12
Board Recommendation			13
Audit Committee Report			14
EXECUTIVE OFFICERS			15
CORPORATE GOVERNANCE			17
Corporate Governance Guidelines			17
Board Leadership Structure			17
Director Independence			17
Board Committees			18
Board of Directors and Committee Meetings and Attendance			20
Executive Sessions			20
Director Attendance at Annual Meeting of Stockholders			20
Director Nominations Process			21
Director Succession Planning and Annual Evaluations			22
Board Role in Risk Oversight			22
Committee Charters and Corporate Governance Guidelines			22
Justice, Equity, Diversity and Inclusion			23
Code of Business Conduct and Ethics			23
Anti-Hedging Policy			23
Stockholder Communications with Our Board of Directors			23
EXECUTIVE OFFICER AND DIRECTOR COMPENSATION			24


Executive Officer Compensation Overview			24
202 1 Summary Compensation Table			24
Narrative Disclosure to 202 1 Summary Compensation Table			25
Outstanding Equity Awards at 202 1 Fiscal Year -End			27
Director Compensation			27
STOCK OWNERSHIP			31
Security Ownership of Certain Beneficial Owners and Management			31
TRANSACTIONS WITH RELATED PERSONS			33
Policies and Procedures Regarding Transactions with Related Persons			33
Certain Related Person Transactions			33
STOCKHOLDER PROPOSALS AND DIRECTOR NOMINATIONS			34
HOUSEHOLDING OF PROXY MATERIALS			35
202 1 ANNUAL REPORT			36


Class I Director - Current Term Ending at 2022 Annual Meeting	Class II Director - Current Term Ending at 2023 Annual Meeting	Class III Director - Current Term Ending at 2024 Annual Meeting
Jose-Carlos Gutiérrez-Ramos, Ph.D.	Juan Andres	David R. Epstein
Iain B. McInnes, M.B.Ch.B., Ph.D.	Lord Ara Darzi	Balkrishan (Simba) Gill, Ph.D.
Theodose Melas-Kyriazi	Julie H. McHugh	John A. Hohneker, M.D.
David P. Perry*		Tonya Williams


Class I Directors	Age	Director Since	Current Position at Evelo
Jose-Carlos Gutiérrez-Ramos, Ph.D.	59	April 2022*	Director
Iain B. McInnes, M.B.Ch.B., Ph.D.	57	September 2021	Director
Theodose Melas-Kyriazi	62	February 2017	Director


Class II Directors	Age	Director Since	Current Position at Evelo
Juan Andres	57	December 2019	Director
Lord Ara Darzi	61	February 2018	Director*
Julie H. McHugh	57	April 2021	Director


Class III Directors	Age	Director Since	Current Position at Evelo
David R. Epstein	60	March 2017	Chair of the Board*
Balkrishan (Simba) Gill, Ph.D.	58	September 2015	Chief Executive Officer, President and Director
John A. Hohneker, M.D.	62	January 2021	Director
Tonya Williams	48	April 2022	Director


	Female	Male	Non-Binary	Did not Disclose Gender
Directors	2	9	—	—
Number of Directors who identify in Any of the Categories Below:
African American or Black	1	—	—	—
Alaskan Native or Native American	—	—	—	—
Asian	—	1	—	—
Hispanic or Latinx	—	1	—	—
Native Hawaiian or Pacific Islander	—	—	—	—
White	1	7	—	—
Two or More Races or Ethnicities	—	—	—	—
LGBTQ+	1
Did Not Disclose Demographic Background	—


	Fiscal Year 2021		Fiscal Year 2020
Audit Fees(1)	$	682,800	$	619,600
Audit-related fees(2)	$	123,500	$	47,000
Tax fees(3)	$	—	$	—
All other fees	$	960	$	2,000
Total Fees	$	807,260	$	668,600


- Board size, independence and qualifications - Executive sessions of independent directors - Board leadership structure - Selection of new directors - Director orientation and continuing education - Limits on board service - Change of principal occupation - Term limits - Director responsibilities - Director compensation						- Stock ownership - Board access to senior management - Board access to independent advisors - Board self-evaluations - Board meetings - Meeting attendance by directors and non-directors - Meeting materials - Board committees, responsibilities and independence - Succession planning - Risk management


Name and Principal Position	Year	Salary ($)	Bonus ($)(1)	Stock Awards ($)(2)	Option Awards ($)(3)	Non-equity incentive plan compensation ($)(4)	All other compensation ($)	Total ($)
Balkrishan (Simba) Gill, Ph.D.	2021	575,000	—	—	3,372,930	268,813	509(5)	4,217,252
Chief Executive Officer	2020	560,000	—	—	1,208,125	400,400	—	2,168,525

Luca Scavo	2021	415,000	50,000	64,721	3,184,290	82,250	98,512(6)	3,894,773
Chief Financial Officer

Jonathan Zung, Ph.D.	2021	415,000	110,000	166,500	3,679,560	141,100	47,177(7)	4,559,336
Chief Development Officer


		Option Awards						Stock Awards
Name and Principal Position	Vesting Start Date	Number of Securities Underlying Unexercised Options (#) Exercisable	Number of Securities Underlying Unexercised Options (#) Unexercisable		Equity Incentive Plan Awards: Number of Securities Underlying Unexercised Unearned Options (#)	Option Exercise Price ($/Sh)	Option Expiration Date	Number of Shares or Units of Stock That Have Not Vested (#)		Market Value of Shares or Units of Stock That Have Not Vested ($) (1)
Balkrishan (Simba) Gill, Ph.D.	7/1/2015	245,854	—		—	0.49	11/4/2025	—		—
Chief Executive Officer	7/1/2016	596,959	—		—	1.14	10/4/2026	—		—
	7/1/2017	183,868	—	(2)	—	3.96	12/14/2027	—		—
	3/21/2018	126,408	8,428	(2)	—	10.48	4/3/2028	—		—
	4/4/2018	—	144,643	(3)	—	10.48	4/3/2028	—		—
	5/8/2018	127,635	18,234	(2)	—	16.00	5/8/2028	—		—
	1/14/2019	151,250	68,750	(2)	—	13.71	1/14/2029	—		—
	1/28/2020	119,791	130,209	(4)	—	7.06	1/27/2030	—		—
	1/27/2021	63,019	211,981	(4)	—	16.65	1/26/2031	—		—

Luca Scavo	6/1/2021	—	300,000	(5)	—	14.24	5/31/2031	—		—
Chief Financial Officer	6/1/2021	—	—		—	—	—	4,545	(6)	$	27,588

Jonathan Zung, Ph.D.	1/4/2021	—	300,000	(5)	—	16.65	1/26/2031	—		—
Chief Development Officer	1/27/2021	—	—		—	—	—	10,000	(7)	$	60,700


Name	Age	Position(s)	In Current Position Since
Balkrishan (Simba) Gill, Ph.D.	58	Chief Executive Officer, President and Director	2015
Mark Bodmer, Ph.D.	67	Chief Scientific Officer and President of Research and Development	2016
Daniel S. Char	61	General Counsel and Secretary	2018
Duncan McHale, M.B.B.S., Ph.D.	55	Chief Medical Officer	2018
Mark D. Plinio	55	Chief Commercial Officer	2021
Luca Scavo	55	Chief Financial Officer, Senior Vice President and Treasurer	2021
Jonathan Zung, Ph.D.	57	Chief Development Officer	2021


	Fees earned or paid in cash		Stock awards		Option awards	All other compensation		Total
Name(1)	($)		($)		($)(2)	($)		($)
Juan Andres	47,958		—		182,942	—		230,900
Lord Ara Darzi	49,417		—		182,942	—		232,359
David R. Epstein	75,000	(3)	812,158	(4)	—	150,000	(5)	1,037,158
Jose-Carlos Gutiérrez-Ramos, Ph.D.	—		—		—	—		—
John Hohneker, M.D.	43,841		—		384,882	—		428,723
Julie McHugh	39,669		—		230,075	8,750	(6)	278,494
Iain B. McInnes, M.B.Ch.B., Ph.D.	15,867		—		286,318	—		302,185
Theodose Melas-Kyriazi	57,917		—		182,942	—		240,859
David P. Perry	50,283		—		183,486	—		233,769
Nancy A. Simonian, M.D.(7)	21,056		—		—	—		21,056
Tonya Williams	—		—		—	—		—


Name(1)						Shares Subject to Options Outstanding at Year-End (#)
Juan Andres						64,570
Lord Ara Darzi						112,621
David R. Epstein						367,891
Jose-Carlos Gutiérrez-Ramos, Ph.D.						—
John Hohneker, M.D.						31,380
Julie H. McHugh						43,148
Iain B. McInnes, M.B.Ch.B., Ph.D.						35,000
Theodose Melas-Kyriazi						105,266
David P. Perry						120,423
Nancy A. Simonian, M.D.(2)						—
Tonya Williams						—


	Shares of Common Stock Beneficially Owned	Percentage of Shares Beneficially Owned
Holders of More than 5%:
Entities affiliated with Flagship Pioneering (1)	23,030,069	42.9	%
Entities affiliated with FMR LLC (2)	8,018,922	14.9	%
Credit Suisse AG (3)	3,566,487	6.6	%
Entities affiliated with HarbourVest Partners (4)	3,063,505	5.7	%
DNCA Finance S.C.S. (5)	2,688,074	5.0	%
Directors and Named Executive Officers:
Balkrishan (Simba) Gill, Ph.D. (6)	1,859,566	3.5	%
Luca Scavo (7)	80,000	*
Jonathan Zung, Ph.D. (8)	115,830	*
Juan Andres (9)	59,340	*
Lord Ara Darzi (10)	112,621	*
David R. Epstein (11)	423,105	*
Jose-Carlos Gutiérrez-Ramos, Ph.D. (12)	12,222	*
John A. Hohneker, M.D. (13)	13,946	*
Julie H. McHugh (14)	23,971	*
Iain B. McInnes, M.B.Ch.B., Ph.D. (15)	8,750	*
Theodose Melas-Kyriazi (16)	105,266	*
David P. Perry (17)	120,423	*
Tonya Williams (18)	2,222	*
All executive officers and directors as a group (17 persons) (19)	4,397,590	8.2	%


*			Represents beneficial ownership of less than 1% of our outstanding common stock.


Participants	Shares of Common Stock	Aggregate Value
Holders of More than 5% (1)
Entities affiliated with Flagship Pioneering	4,666,666	$	17,499,998
Entities affiliated with FMR LLC	1,466,666	$	5,499,998
Entities affiliated with Harbour Vest Partners	1,300,000	$	4,875,000