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NOTICE OF ANNUAL MEETING OF STOCKHOLDERS

PROXY STATEMENT		1

Proposals		1
Recommendations of the Board		1
Information About This Proxy Statement		2

QUESTIONS AND ANSWERS ABOUT THE 2021 ANNUAL MEETING OF STOCKHOLDERS		3

PROPOSALS TO BE VOTED ON		7

Proposal 1: Election of Directors		7
Proposal 2: Ratification of Appointment of Independent Registered Public Accounting Firm		11

REPORT OF THE AUDIT COMMITTEE OF THE BOARD OF DIRECTORS		12

INDEPENDENT REGISTERED PUBLIC ACCOUNTING FIRM FEES AND OTHER MATTERS		13

EXECUTIVE OFFICERS		14

CORPORATE GOVERNANCE		15

General		15
Board Composition		15
Director Independence		15
Director Candidates		15
Communications from Stockholders		16
Board Leadership Structure and Role in Risk Oversight		16
Code of Ethics		17
Anti-Hedging Policy		17
Attendance by Members of the Board of Directors at Meetings		17

COMMITTEES OF THE BOARD		18

Audit Committee		18
Compensation Committee		19
Nominating and Corporate Governance Committee		19

EXECUTIVE AND DIRECTOR COMPENSATION		20

Executive Compensation		20
2020 Summary Compensation Table		20
Narrative Disclosure to Summary Compensation Table		20
Outstanding Equity Awards at 2020 Fiscal Year End		22
Employment Agreements		22
Non-Employee Director Compensation		23
2020 Director Compensation Table		23

SECURITY OWNERSHIP OF CERTAIN BENEFICIAL OWNERS AND MANAGEMENT		25

CERTAIN RELATIONSHIPS AND RELATED PERSON TRANSACTIONS		27

STOCKHOLDERS’ PROPOSALS		28

OTHER MATTERS		29

SOLICITATION OF PROXIES		29

HOMOLOGY’S ANNUAL REPORT ON FORM 10-K		29


	•		by Internet—You can vote over the Internet at www.proxyvote.com by following the instructions on the Internet Notice or proxy card;


	•		by Telephone—You can vote by telephone by calling 1-800-690-6903 and following the instructions on the Internet Notice or proxy card;


	•		by Mail—You can vote by mail by signing, dating and mailing the proxy card, which you may have received by mail; or

	•		Electronically at the Meeting—If you attend the meeting online, you will need the 16-digit control number included in your Internet Notice, on your proxy card or on the instructions that accompanied your proxy materials to vote electronically during the meeting.


	•		by submitting a duly executed proxy bearing a later date;


	•		by granting a subsequent proxy through the Internet or telephone;


	•		by giving written notice of revocation to the Secretary of Homology prior to or at the Annual Meeting; or


Proposal	Votes required	Effect of Votes Withheld / Abstentions and Broker Non-Votes
Proposal 1: Election of Directors	The plurality of the votes cast. This means that the three nominees receiving the highest number of affirmative “FOR” votes will be elected as Class III Directors.	Votes withheld and broker non-votes will have no effect.

Proposal 2: Ratification of Appointment of Independent Registered Public Accounting Firm	The affirmative vote of the holders of a majority in voting power of the votes cast affirmatively or negatively.	Abstentions will have no effect. We do not expect any broker non-votes on this proposal.


		The Board of Directors unanimously recommends a vote FOR the election of the below Class III Director nominees.

Name	Age	Position with Homology	Served as a Director Since
Matthew R. Patterson	49	Director	2018
Mary Thistle	61	Director	2018
Arthur O. Tzianabos, Ph.D.	58	President and Chief Executive Officer and Director	2016

	2020		2019
Fee Category	(in thousands)
Audit Fees	$	598	$	602
Tax Fees		20		13
All Other Fees		2		28
Total	$	620	$	643

Name	Age	Position
Arthur O. Tzianabos, Ph.D. (1)	58	President and Chief Executive Officer and Director
W. Bradford Smith (2)	65	Chief Financial Officer and Treasurer
Albert Seymour, Ph.D. (3)	53	Chief Scientific Officer
Tim Kelly (4)	52	Chief Operating Officer
Gabriel M. Cohn, M.D. (5)	62	Chief Medical Officer

(1)	See biography on page 8 of this proxy statement.
(2)	W. Bradford Smith has served as our Chief Financial Officer and Treasurer since April 2017 and also served as our Secretary from July 2017 to June 2020. From March 2014 to April 2017, Mr. Smith was Chief Financial Officer of Ocular Therapeutix, Inc., a biopharmaceutical platform company. Prior to joining Ocular Therapeutix, Inc., Mr. Smith served as the Chief Financial Officer of OmniGuide, Inc., a medical device company, from July 2008 to March 2014. Mr. Smith holds a B.S. in Biology from Tufts University and an M.B.A. from the Whittemore School of Business and Economics at the University of New Hampshire.
(3)	Albert Seymour, Ph.D. has served as our Chief Scientific Officer since April 2016. Prior to joining Homology, Dr. Seymour was Senior Vice President, Head of Global Research and Nonclinical Development at Shire plc, a biotechnology company, from 2011 to 2016. Dr. Seymour received his B.A. in Biology from the University of Delaware, an M.S. from Johns Hopkins University School of Medicine and his Ph.D. in Human Genetics from the University of Pittsburgh.
(4)	Tim Kelly has served as our Chief Operating Officer since April 2021, and prior to that, he served as our Chief Technical Operations Officer from April 2018 to April 2021, and as our Senior Vice President of Technical Operations from May 2017 to April 2018. From January 2017 to May 2017, Mr. Kelly served as Head of Technical Operations at Sarepta Therapeutics, Inc., a biopharmaceutical company. Prior to that, Mr. Kelly spent eight years, from 2009 to 2017, at Shire plc, a biotechnology company, where he served most recently as Senior Vice President and Head of Biologics Operating Unit from May 2016 to January 2017, and prior to that served in roles of increasing responsibility, including Interim Head of Technical Operations from August 2015 to April 2016 and Head of Product Strategy & Planning from 2014 to July 2015. He was previously the Head of Technical Operations at UCB S.A. in Belgium. Mr. Kelly also held roles of increasing responsibility at Biogen Idec, including Senior Director of International Operations. Mr. Kelly holds a B.S. with emphasis in Engineering Mechanics from the United States Air Force Academy and an M.B.A. from Troy University.
(5)	Gabriel M. Cohn, M.D. has served as our Chief Medical Officer since December 2019. Prior to joining Homology, Dr. Cohn was Vice President, Clinical Development Lead at AVROBIO, Inc., a clinical-stage gene therapy company, from November 2017 to November 2019. Prior to AVROBIO, Dr. Cohn served as Vice President and Head of Global Clinical and Medical Affairs at OvaScience, Inc., a biotechnology company, from 2015 to July 2017. Prior to that, Dr. Cohn spent seven years at Shire Plc, a biotechnology company, including Shire Human Genetic Therapies (HGT), where he most recently held the position of Senior Medical Director, Clinical Sciences from 2012 to 2015, and prior to that served in roles of increasing responsibility including Interim Global Franchise Medical Lead, Gaucher Disease, in 2011 and Medical Director, North America, from 2008 to 2011. Prior to joining the biotechnology industry, Dr. Cohn served as the Chief, Division of Clinical and Reproductive Genetics and Medical Director, Genetic Services at Baystate Medical Center and as Assistant Professor at Tufts University School of Medicine. He began his academic career as the Director, Reproductive Genetics at SUNY Health Science Center (HSC) at Syracuse (now Upstate). Dr. Cohn has a B.S. in Biology from Brooklyn College, an M.D. from SUNY HSC at Syracuse School of Medicine and an M.B.A. from University of Massachusetts Amherst.



Name	Audit	Compensation	Nominating and Corporate Governance
Steven Gillis, Ph.D.			Chairperson
Richard J. Gregory, Ph.D.		X
Kush M. Parmar, M.D., Ph.D.		Chairperson
Matthew R. Patterson	X
Jeffrey V. Poulton	X
Alise Reicin, M.D.			X
Mary Thistle	Chairperson

	•		appointing, approving the compensation of, and assessing the independence of our independent registered public accounting firm;
	•		overseeing the work of our independent registered public accounting firm, including through the receipt and consideration of reports from such firm;
	•		reviewing and discussing with management and the independent registered public accounting firm our annual and quarterly financial statements and related disclosures;
	•		monitoring our internal control over financial reporting, disclosure controls and procedures and code of business conduct and ethics;
	•		discussing our risk management policies;
	•		establishing policies regarding hiring employees from the independent registered public accounting firm and procedures for the receipt and retention of accounting-related complaints and concerns;
	•		meeting independently with our internal auditing staff, if any, independent registered public accounting firm and management;
	•		reviewing and approving or ratifying any related person transactions; and
	•		preparing the audit committee report required by the SEC rules (which is included on page 12 of this proxy statement).


	•		reviewing and approving, or recommending for approval by the Board, the compensation of our CEO and our other executive officers;
	•		overseeing and administering our cash and equity incentive plans;
	•		reviewing and making recommendations to the Board of Directors with respect to director compensation;
	•		reviewing and discussing annually with management our “Compensation Discussion and Analysis,” to the extent required; and
	•		preparing the annual compensation committee report, to the extent required by SEC rules.


	•		identifying individuals qualified to become board members;
	•		recommending to the Board of Directors the persons to be nominated for election as directors and to each board committee;
	•		reviewing the Board’s leadership structure;
	•		developing and recommending to the Board of Directors corporate governance guidelines; and
	•		overseeing an annual evaluation of the Board of Directors.


	•		Arthur O. Tzianabos, Ph.D., President and Chief Executive Officer;
	•		Albert Seymour, Ph.D., Chief Scientific Officer; and
	•		W. Bradford Smith, Chief Financial Officer.

Name and principal position	Fiscal Year	Salary $		Bonus $		Option Awards $ (1)		Non-Equity Incentive Plan Compensation $		All Other Compensation $ (2)		Total $
Arthur O. Tzianabos, Ph.D.	2020		563,200		—		—		309,760		13,000		885,960
President and Chief Executive Officer	2019		537,600		—		4,217,097		295,680		12,500		5,062,877
Albert Seymour, Ph.D.	2020		426,800		—		—		170,720		10,057		607,577
Chief Scientific Officer	2019		405,600		—		1,295,756		162,240		13,231		1,876,827
W. Bradford Smith	2020		406,700		—		—		162,680		13,000		582,380
Chief Financial Officer	2019		384,200		—		1,307,536		153,680		14,880		1,860,296

Name	2020 Annual Base Salary ($)		2021 Annual Base Salary ($)
Arthur O. Tzianabos, Ph.D.		563,200		583,100
Albert Seymour, Ph.D.		426,800		450,000
W. Bradford Smith		406,700		445,300

	Option Awards
Name	Vesting Commencement Date	Number of Securities Underlying Unexercised Options (#) Exercisable		(1)	Number of Securities Underlying Unexercised Options (#) Unexercisable		(1)	Per Share Option Exercise Price ($)		Option Expiration Date
Arthur O. Tzianabos, Ph.D.	4/1/2016		462,135	(2)		—	(2)		0.47	4/22/2026
	1/1/2018		375,242			139,411			6.63	12/7/2027
	3/27/2018		63,492			28,884			16.00	3/27/2028
	1/1/2019		97,270			105,730			24.28	12/14/2028
	1/1/2020		82,041			275,959			19.92	12/11/2029
Albert Seymour, Ph.D.	1/1/2018		55,702			20,715			6.63	12/7/2027
	3/27/2018		-			4,176			16.00	3/27/2028
	1/1/2019		35,458			38,542			24.28	12/14/2028
	1/1/2020		25,208			84,792			19.92	12/11/2029
W. Bradford Smith	4/5/2017		89,786	(2)		13,618	(2)		0.63	4/6/2027
	1/1/2018		46,283			17,213			6.63	12/7/2027
	3/27/2018		34,155			15,556			16.00	3/27/2028
	1/1/2019		42,166			45,834			24.28	12/14/2028
	1/1/2020		25,437			85,563			19.92	12/11/2029

Name	Fees Earned or Paid in Cash ($)		Option Awards ($)(1)			Total ($)
Steven Gillis, Ph.D.		48,000		164,759	(2)		212,759
Richard J. Gregory, Ph.D.		48,750		164,759	(2)		213,509
Kush M. Parmar, M.D., Ph.D.		85,000		164,759	(2)		249,759
Matthew R. Patterson		47,500		164,759	(2)		212,259
Jeffrey V. Poulton (3)		23,750		342,900	(4)		366,650
Alise S. Reicin, M.D.		44,000		164,759	(2)		208,759
Mary Thistle		55,000		164,759	(2)		219,759

(1)	Amounts reflect the full grant date fair value of stock options granted during the applicable year computed in accordance with ASC Topic 718, rather than the amounts paid to or realized by the named individual. We provide information regarding the assumptions used to calculate the value of all option awards in Note 12 to our consolidated financial statements included in our Annual Report on Form 10-K for the fiscal year ended December 31, 2020.
(2)	Represents the grant date fair value of an option to purchase 18,000 shares of our common stock granted to each then-current non-employee director on June 10, 2020 with an exercise price of $16.32 per share.
(3)	Mr. Poulton was appointed to our Board of Directors on July 21, 2020. Mr. Poulton’s fees for his board and committee service were prorated as if he had served during the entire second half of 2020.
(4)	Represents the grant date fair value of an initial grant of an option to purchase 36,000 shares of our common stock with an exercise price of $16.48 per share granted to Mr. Poulton upon his appointment as a director on July 21, 2020.

	Options Outstanding
Name	Exercisable		Unexercisable		Total
Steven Gillis, Ph.D.		36,145		28,595		64,740
Richard J. Gregory, Ph.D.		36,145		28,595		64,740
Kush M. Parmar, M.D., Ph.D.		36,145		28,595		64,740
Matthew R. Patterson		38,411		27,279		65,690
Jeffrey V. Poulton (3)		—		36,000		36,000
Alise S. Reicin, M.D.		10,282		38,878		49,160
Mary Thistle		36,145		28,595		64,740

Name of Beneficial Owner	Number of Shares Beneficially Owned		Percentage
5% or Greater Stockholders
Entities affiliated with ARCH Venture Fund (1)		5,768,694		10.1	%
Pfizer Inc. (2)		5,000,000		8.8	%
Entities affiliated with 5AM Ventures (3)		4,535,919		7.9	%
JPMorgan Chase & Co. (4)		3,441,712		6.0	%
TLS Beta Pte. Ltd. (5)		3,220,293		5.6	%
Named Executive Officers and Directors
Arthur O. Tzianabos, Ph.D. (6)		1,328,225		2.3	%
Albert Seymour, Ph.D. (7)		338,697	*
W. Bradford Smith (8)		288,647	*
Steven Gillis, Ph.D. (1)(9)		5,833,434		10.2	%
Richard J. Gregory, Ph.D. (10)		75,546	*
Kush M. Parmar, M.D., Ph.D. (3)(11)		4,600,659		8.1	%
Matthew R. Patterson (12)		61,729	*
Jeffrey V. Poulton		—	*
Alise S. Reicin, M.D. (13)		28,282	*
Mary Thistle (14)		64,740	*
All executive officers and directors as a group (12 persons) (15)		12,905,484		21.7	%


*	Less than one percent.
(1)	Based on a Schedule 13G/A filed with the SEC on February 13, 2020 and the Company's records. Consists of 4,631,031 shares of common stock held by ARCH Venture Fund VIII, L.P. (“ARCH Fund VIII”) and 1,137,663 shares of common stock held by ARCH Venture Fund VIII Overage, L.P. (“ARCH Fund Overage”). The sole general partner of ARCH Fund VIII is ARCH Venture Partners VIII, L.P. (“ARCH Partners VIII”), which may be deemed to beneficially own the shares held by ARCH Fund VIII. The sole general partner of ARCH Partners VIII and ARCH Fund Overage is ARCH Venture Partners VIII, LLC (“ARCH VIII LLC”), which has shared voting and dispositive power over the shares of common stock held by each of ARCH Fund VIII and ARCH Fund Overage. ARCH Partners VIII and ARCH VIII LLC disclaim beneficial ownership of such shares, except to the extent of any pecuniary interest therein. The managing directors of ARCH VIII LLC are Keith L. Crandell, Clinton Bybee and Robert Nelsen, and they may be deemed to have shared voting and dispositive power over the shares of common stock held by ARCH Fund VIII and ARCH Fund Overage. Messrs. Crandell, Bybee and Nelsen disclaim beneficial ownership of such shares, except to the extent of any pecuniary interest therein. Steven Gillis, M.D., Ph.D., one of our directors, is a managing director at ARCH Venture Partners. Director Steven Gillis owns an interest in ARCH Partners VIII but does not have voting or investment control over the shares held by ARCH Fund VIII, and disclaims beneficial ownership of such shares, except to the extent of any pecuniary interest therein. The address of ARCH Fund VIII and ARCH Fund Overage is 8755 West Higgins Road, Suite 1025, Chicago, Illinois 60631.
(2)	Based on a Schedule 13G filed with the SEC on November 19, 2020, Pfizer Inc. has sole voting and dispositive power over all 5,000,000 shares. The address of Pfizer Inc. is 235 East 42nd Street, New York, NY 10017.

(3)	Based on a Schedule 13G/A filed with the SEC on February 16, 2021 and the Company's records. Consists of 4,354,484 shares of common stock held by 5AM Ventures IV, L.P. (“Ventures IV”), as to which Ventures IV has shared voting and dispositive power, and 181,435 shares of common stock held by 5AM Co-Investors IV, L.P. (“Co-Investors IV”), as to which Co-Investors IV has shared voting and dispositive power. 5AM Partners IV, LLC (“Partners IV”) is the sole general partner of Ventures IV and Co-Investors IV. Dr. John Diekman, Andrew J. Schwab and Dr. Scott M. Rocklage, are the managing members of Partners IV and, along with Partners IV, have shared voting and investment power over the shares beneficially owned by Ventures IV and Co-Investors IV. Kush M. Parmar, M.D., Ph.D., one of our directors, is an affiliate of Ventures IV. Each of Partners IV, Dr. Diekman, Mr. Schwab and Dr. Rocklage disclaim beneficial ownership of such shares except to the extent of its or their recurring interest therein. The address of all entities affiliated with 5AM Ventures is 501 2nd Street, Suite 350, San Francisco, CA 94107.
(4)	Based solely on a Schedule 13G/A filed with the SEC on January 25, 2021, JPMorgan Chase & Co. has sole voting power over 3,054,192 shares and sole dispositive power over all 3,441,712 shares. The address of JPMorgan Chase & Co is 383 Madison Avenue, New York, NY 10179.
(5)	Based solely on a Schedule 13G filed with the SEC on April 11, 2018, Temasek Holdings (Private) Limited, Fullerton Management Pte Ltd and Temasek Life Sciences Private Limited each has shared voting and dispositive power over 3,220,293 shares of common stock, V-Sciences Investments Pte Ltd has shared voting and dispositive power over 625,000 shares of common stock, and TLS Beta Pte. Ltd. has shared voting and dispositive power over 2,595,293 shares of common stock. The principal business address of Temasek Holdings (Private) Limited, Fullerton Management Pte Ltd, Temasek Life Sciences Private Limited, V-Sciences Investments Pte Ltd and TLS Beta Pte. Ltd. is 60B Orchard Road #06-18 Tower 2, The Atrium@Orchard, Singapore 238891.
(6)	Includes options to purchase 1,250,720 shares of common stock that are or will be immediately exercisable by Dr. Tzianabos within 60 days of April 23, 2021.
(7)	Includes options to purchase 209,680 shares of common stock that are or will be immediately exercisable by Dr. Seymour within 60 days of April 23, 2021.
(8)	Consists of options to purchase 288,647 shares of common stock that are or will be immediately exercisable by Mr. Smith within 60 days of April 23, 2021.
(9)	Includes options to purchase 64,740 shares of common stock that are or will be immediately exercisable by Dr. Gillis within 60 days of April 23, 2021.
(10)	Includes options to purchase 64,740 shares of common stock that are or will be immediately exercisable by Dr. Gregory within 60 days of April 23, 2021.
(11)	Includes options to purchase 64,740 shares of common stock that are or will be immediately exercisable by Dr. Parmar within 60 days of April 23, 2021.
(12)	Consists of options to purchase 61,729 shares of common stock that are or will be immediately exercisable by Mr. Patterson within 60 days of April 23, 2021.
(13)	Consists of options to purchase 28,282 shares of common stock that are or will be immediately exercisable by Ms. Reicin within 60 days of April 23, 2021.
(14)	Consists of options to purchase 64,740 shares of common stock that are or will be immediately exercisable by Ms. Thistle within 60 days of April 23, 2021.
(15)	Includes options to purchase 2,383,543 shares of common stock that are or will be immediately exercisable within 60 days of April 23, 2021.


(1)	Affiliate of Cameron Wheeler, Ph.D., who resigned from our Board of Directors in July 2019.